For the most part, harpacticoids listed below were taken from the taxa found by Lang (1965), Kask et al. (1982), Watkins (1983), Webb (1989), and from collections made by the author in Puget Sound and in eelgrass beds and coastal estuaries of Washington, Oregon, and California. Information for the annotations also came from these sources, from Lang (1948), and from the author's observations; it should be noted that many habitats remain unsampled and any species may occur in more habitats/regions than listed. In many cases, the only record for a species is from Lang's (1965) original collections, mainly from near Hopkins Marine Station, Pacific Grove (abbreviated as HML in the species list, below), and Pacific Marine Station, Dillon Beach, California (DB). Chappuis recorded 76 (mainly european) species of harpacticoids from Puget Sound, Washington, but Lang (1965) refuted most of these, and they are not included in the list. Similarly, Crandell (1966) listed 57 mainly northern european species in Yaquina Bay, Oregon that were identified just prior to Lang's (1965) regional monograph, and are also not listed unless also recorded elsewhere within the region. Taxa designated with an asterisk have not been recorded from California or Oregon, but occur in Washington and/or southern B.C.. Also listed are several species of algae-dwelling harpacticoids that were found near La Jolla, in southern California (Gunnill 1982). Indicates that additional undescribed or unidentified species have been observed by the author, for that genus, within the region covered. This designation includes species in the genera Tegastes, Dactylopusia, Psyllocamptus, Mesochra, and Itunella that were described in a PhD thesis (Watkins 1983) but have not yet been published in refereed journals. Lang (1948) summarized harpacticoid taxonomic literature, and gave keys to families, genera, and species. Bodin (1997) gives a complete listing of all additional descriptions, new species, revisions, and taxonomic publications that have occurred since. Huys et al. (1996) give an up-to-date key to the marine and brackish water harpacticoid families. Where keys published subsequent to Lang's 1948 monograph exist, they are cited for the families and genera listed.
Coullana canadensis (Willey, 1923). Usually dominant epibenthic and planktonic copepod in oligohaline to tidal fresh reaches of large rivers; also common in smaller estuaries. Description of male and redescription of female (as Scottolana canadensis ): Coull 1972, Crustaceana 22(3): 209-214.
Longipedia americana Wells, 1980. Lower intertidal into the subtidal, in medium sand and/or mixtures of sand and mud. Abundant in sediment around eelgrass, Zostera marina, and often occurs in near-bottom plankton net samples. Other species in this genus may be present. Key to the species: Wells 1980, Zool. J. Linn. Soc. 70: 103-189..
Huys 1990 (see below) formed this family from several species previously assigned to the genus Rhynchothalestris (family Thalestridae).
Ambunguipes rufocincta (Brady, 1880)*. Uncommon on eelgrass and macroalgae in Puget Sound and Hood Canal, WA. Com. nov. for Rhynchothalestris rufocincta (Brady, 1880). See Huys 1990, Zool. J. Linn. Soc. 99: 51-115.
Key to the genera: Lang 1965, Petkovski 1976, Acta Mus. Maced. Sci. Nat. 15 (1): 1-26..
Ameira longipes Boeck, 1864 . Geographically widespread and common in high intertidal pools to shallow subtidal eelgrass beds where microalgal epiphytes and detritus occur. Redescription: Lang (1965) from near HMS.
A. parvuloides Lang, 1965.
A. parasimulans Lang, 1965. This and previous species uncommon in collections from tidal pools and eelgrass; may be affiliated with sediment. Described from interstitial shell and sand at DB.
Ameira minuta Boeck, 1865. Sediments and leaves in eelgrass and subtidal mud and gravel.
Ameiropsis longicornis Sars, 1907*. North Atlantic and Mediterranean species found in shallow subtidal sand in southern B.C. (Kask et al. 1982).
Interleptomesochra reducta Lang, 1965. Interstitial fine shell sand. Key to the species: Lang 1965.
Nitokra typica Boeck, 1865 . Eelgrass and sandy mud in estuaries.
N. spinipes Boeck, 1865. Estuarine marshes and tidepools with freshwater input. Lang (1965) described the CA variant of this widespread species as N. s. armata, but Wells & Rao [1987, Mem. Zool. Surv. India, 16 (4): 1-385] rejected subspecies of N. spinipes.
N. lacustris (Schmankevitch, 1875). Low-salinity and fresh coastal and inland marshes.
Nitokra affinis californica Lang, 1965. Described from a tide pool with shell sand, stones, and algae near HMS.
Praeleptomesochra similis Lang, 1965. Tidal pools, shell sand, stones, algae. Key to the species: Lang 1965, Pesce 1981, Bull. Zool. Mus. Univ. Amsterdam, 8 (9): 69-72..
Pseudoleptomesochra typica Lang, 1965. Interstitial shell sand.
Psyllocamptus triarticulatus Lang, 1965 . This and previous species described from tidal pools containing shell sand, stones, and algae off HMS. Watkins (1983) collected another species in Mendocino and San Luis Obispo counties, CA. Key to the species: Lang 1965, Ceccherelli 1988, Vie Milieu 38 (2): 155-171..
Psyllocamptus minutus Sars, 1911*. Shallow subtidal sand and cobble.
Sarsameira sp.*. Intertidal mud in estuaries. Key to the species: Reidenauer & Thistle 1983, Trans. Am. Micros. Soc. 102 (2): 105-112.
Family Ancorabolidae Key to genera: Lang 1965
Laophontodes hedgepethi Lang, 1965. Associated with Laminaria and other algal holdfasts. In New England, this and other genera in the family also associated with colonial invertebrates.
See Bodin (1997) for notes on revisions of this family.
Itunella muelleri (Gagern, 1922) . European species reported from the Santa Rosa creek estuary by Watkins (1983), who also reported an undescribed species from supralittoral pools.
Mesochra pygmaea (Claus, 1863) . Usually affiliated with vegetation; one of the most abundant and widespread harpacticoids in eelgrass along the Pacific coast; also recorded from brackish water. Key to the species: Hamond 1971, Austral. J. Zool. suppl. 7: 1-32, Fiers & Rutledge 1990, Bull. Inst. R. Sci. Nat. Belg., Biol. 60: 105-125.
Mesochra rapiens (Schmeil, 1894)*. Mud and sand in oligohaline and tidal freshwater reaches of estuaries.
Mesochra alaskana M.S. Wilson, 1958*. Same as previous species, in more saline regions.
Nannomesochra arupinensis (Brian, 1925). Widely distributed in warm marine and brackish waters; in CA found in Enteromorpha in brackish pools (Watkins 1983).
Leimia vaga Willey, 1923*. Benthic and epibenthic; often abundant in estuarine mudflats and tidal channels.
Cletocamptu deitersi Schmankevitch, 1875. Cosmpopolitan in coastal brackish and inland saline environments, including desert springs in CA (Watkins 1976). Revision of the genus: Fleeger 1980, Trans. Am. Micros. Soc. 99 (1): 25-31.
Lang (1965) gives a key to the genera of the former family Cletodidae which encompasses the families Cletodidae s. str., Paranannopidae, Huntemanniidae, Rhizothricidae, Argestidae, and part of the Canthocamptidae.
Acrenydrosoma karlingi Lang, 1965. Tidal pools with coarse sand. Unidentified species which may be A. macalli Schizas and Shirley 1994 [Crustaceana 67 (3): 331-340] occurs in lower intertidal/shallow subtidal mud and fine sand with detritus.
Cletodes hartmannae Lang, 1965. Described from tidal pools with shell sand, stones, and algae near HMS. Key to the species: Lang 1965, Hamond 1973, Mem. Queensland Mus. 16 (3): 471-483, Fiers 1991, Beaufortia 42 (2): 13-47.
Enhydrosoma hopkinsi Lang, 1965. Described from tidal pools with shell sand, stones, and algae near HMS. Key to the species: Thistle 1980, Trans. Am. Microsc. Soc. 99: 384-397, Bell & Kern 1983, Bull. Mar. Sci. 33 (4): 899-904..
Kollerua breviarticulatum Shen and Tai, 1964*. Intertidal and shallow subtidal eelgrass, mud, and sand. Described from freshwater in south China and may be an introduced species. Comb. nov. by Gee 1994 Sarsia, 79: 83-107, for Enhydrosoma breviarticulatum .
Limnocletodes behningi Borutzky, 1926*. Common in estuaries near river mouths, and abundant in intertidal mud near vegetation such as Spartina spp. Revision of the genus: Wells 1971, J. Nat. Hist. 5: 507-520.
Strongylacron buchholzi (Boeck, 1872)*. Mud and sandy mud in estuaries. Comb. nov. by Gee & Huys 1996, Sarsia 81: 161-191 for Enhydrosoma buchholzi.
Stylicletodes verisimilis Lang, 1965. Described from Monterey Bay in fine sand and detritus at about 26 m depth. Key to the species: Bodin 1968, M/span>
Clytemnestra rostrata (Brady 1883).
Clytemnestra scutellata Dana 1848. This and the previous species are planktonic and widespread throughout tropical and temperate oceanic waters down to 60 m, but are sometimes collected near shore.
Key to the genera: Huys et al. 1996
Danielssenia typica Boeck, 1872. Intertidal and shallow subtidal sand and sandy mud; sediment in eelgrass. Some previous records are probably D. reducta . See Gee 1988, Zool. Scr. 17 (1): 39-53 and Huys and Gee 1993, Bull. Nat. Hist. Mus. 59 (1): 45-81 for revisions and keys to the species.
D. reducta Gee 1988*. Sediments around eelgrass.
Key to the genera: Huys et al. 1996
Leptocaris armatus Lang, 1965. Associated with algae and/or algal detritus. Key to the species: Lang 1965, Fleeger & Clark 1980, Northeast Gulf Sci. 3 (2): 53-59..
L. doughertyi Lang, 1965.
L. pori Lang 1965. This and the previous species were described from interstitial fine shell sand in Monterey Bay.
L. sp.*. Kask et al. 1982 found an unidentified Leptocaris species in intertidal sandy mud of several B.C. estuaries. Another, possibly the same species is associated with woody and other plant debris in Puget Sound and coastal estuaries.
Key to the genera: Lang 1965.
Amonardia normani (Brady, 1872)*. Eelgrass and macro- and microalgae. Key to the species (males only): Lang 1965.
A. perturbata Lang 1965. Often very abundant on eelgrass leaves and macroalgae.
Amphiascoides dimorphus Lang 1965 .. Described from intertidal shell sand at DB. Key to the species: Lang 1965.
Amphiascoides lancisetiger Lang 1965. Described from tidal pool with shell sand, stones, and algae near HMS.
Amphiascoides petkovskii Lang, 1965. Intertidal and shallow subtidal sandy mud, including around eelgrass.
Amphiascoides subdebilis (Willey, 1935)*. In sediments around intertidal and subtidal eelgrass and unvegetated sand and mud flats. Nearly cosmopolitan distribution.
Amphiascopsis cinctus (Claus, 1866). Intertidal sand and mud with microalgae, intertidal algae, epiphyton on docks. Cosmopolitan.
Amphiascus minutus (Claus, 1863) . Intertidal eelgrass and sandy mud, and a variety of mid- to low rocky intertidal habitats.
Amphiascus parvus Sars, 1906. European species recorded from at least one site in CA (Watkins 1983) and abundant in eelgrass beds of coastal embayments in OR and WA.
Amphiascus undosus Lang, 1965. Lower intertidal habitats. In OR and WA, an abundant species on eelgrass in estuaries.
Bulbamphiascus imus (Brady, 1872)*. Uncommon in low intertidal/shallow subtidal sand, except in areas of high organic loading or pollution, where it can dominate. Key to the species: Dinet 1971, Tethys 2 (3): 747-762.
Diosaccus spinatus Campbell, 1929 . A prominent bright yellow species on macroalgae and eelgrass. In WA and B.C. this species is rare in the spring, but abundant from late summer into the winter. Key to the species: Lang 1965.
Paramphiascella xiphophora Lang, 1965. Described from off HMS in about 27 m of water on a bottom of fine sand, detritus, and algae. Key to the species: Marcotte 1974, Zool. J. Linn. Soc. 55 (1): 65-82.
Paramphiascopsis ekmani Lang, 1965. Tide pools in mid- to lower intertidal. Key to the species: Lang 1965, Hicks 1986, New Zealand. J. Nat. Hist. 20: 389-397.
Robertgurneya diversa Lang, 1965 .
Robertgurneya hopkinsi Lang, 1965. This and the previous species were described from off HMS at about 26 m depth on a bottom of fine sand, detritus, and algae; both also occur intertidally. Species similar to R. diversa and hopkinsi, as well as several unidentified species have been found in intertidal sand in Puget Sound. Key to the species: Lang 1965, Hamond 1973, J. Nat. Hist. Lond. 7 (1): 65-76.
Robertsonia knoxi (Thompson and A. Scott, 1903). Sheltered sand beaches and lower intertidal/shallow subtidal sand and sandy mud with detritus or eelgrass. Key to the species: Hamond 1973, Rec. Aust. Mus. Sydney 28 (18): 421-435, Fiers 1996, Bull. Mar. Sci. 58 (1): 117-130.
R. propinqua (T. Scott, 1893)*. Abundant in the autumn and winter in subtidal sand and sediments in eelgrass.
Schizopera knabeni Lang, 1965 . Sediments around estuarine eelgrass; extends upstream into oligohaline tidal reaches of estuaries. Co-occurs with one or more unidentified species in Puget Sound. Key to the species: Lang 1965, Apostolov 1973, Acta Mus. Maced. Sc. Nat., Skopje 13 (5): 81-107.
Schizopera sp. Watkins (1983) found an unidentified species in a salt marsh at Chorro creek, San Luis Obispo County, CA.
Stenhelia (Stenhelia) asetosa Thistle and Coull, 1979*. Key to species of the subgenus: Thistle and Coull 1979, Zool. J. Linn. Soc. 66: 63-72.
S. (S.) peniculata Lang, 1965. Described from off HMS, at about 26 m depth on a bottom of fine sand and detritus but is also abundant intertidally. Unlike most of its congeners, this species occurs in algae- and seagrass and rich habitats.
S. (Delavalia) latipes Lang, 1965. Described from same habitat as previous species. Key to the species of the subgenus: Coull 1976, Zool. J. Linn. Soc. 59 (4): 353-364.
S. (D.) latioperculata It™ 1981*. Described from sandy bottom at about 26 m off Hokkaido, Japan. Abundant in intertidal/shallow subtidal sediments near eelgrass and Spartina in WA and B.C..
S. (D.) longicaudata longicaudata Boeck, 1872*. N. Atlantic species found by Kask et al. (1982) in subtidal mud in Departure Bay, B.C..
S. (D.) longipilosa Lang 1965. Described from same habitat as S. (S.) peniculata.
S. (D.) oblonga Lang 1965. Intertidal mud flats and salt marshes.
Typhlamphiascus pectinifer Lang, 1965 Described from Monterey Bay tidal pools containing shell sand, stones, and algae. Occurs in a variety of lower intertidal/shallow subtidal habitats in Puget Sound.
T. unisetosus Lang, 1965. Described from same habitat as the previous species.
Family Ectinosomatidae. Key to the genera: Lang 1965, Huys et al. 1996.
Arenosetella kaiseri Lang, 1965. Described from interstitial shell sand off HMS. Key to the species: Lang 1965, McLachlan & Moore 1978, Ann. S. Afr. Mus., 76 (4): 191-211, Bodin 1979, Vie Milieu, 27 (3-A): 311-357.
Bradya cladiofera Lang, 1965. Described from off HMS in fine sand with detritus, at about 26 m depth.
Bradyellopsis foliatus Watkins, 1987. Collected infrequently by Watkins (1983) by plankton net from lower intertidal/shallow subtidal sites dominated by macroalgae. Key to the species: Watkins 1987, J. Crust. Biol., 7 (2): 380-395.
Ectinosoma melaniceps Boeck, 1865 . Cosmopolitan species; very common in lower intertidal marine habitats rich in detritus and in eelgrass; less common in brackish water. Redescription: Lang 1965, Pallares 1970, Physis, 30 (80): 255-282. Key to the species: Lang 1965.
E. breviarticulatum Lang, 1965.
E. paranormani Lang, 1965. This and the previous species were described from off HMS in a tidal pool with sand and algae.
E. californicum Lang, 1965. Described from rinsing of algae collected off HMS.
Halectinosoma kunzi Lang, 1965 . Described from off HMS in a tidal pool with shell and gravel.
H. longisetosum Lang, 1965. Described from off HMS in a tidal pool with sand and gravel.
H. ornatum Lang, 1965. Described from rinsings of algae from tide pool at Point Pinos, Monterey Bay.
H.similidistinctum Lang, 1965. Described from off HMS in fine sand and detritus at about 26 m depth. Key to the species: Lang 1965, Cloore 1995, Zool. J. Linn. Soc., 114: 247-306.
H. unicum Lang, 1965. Described from off HMS in fine sand with some detritus at about 7 m depth.
Hastigerella abbotti Lang, 1965. Described from intertidal interstitial shell sand off HMS. Key to the species: Lang 1965, McLachlan & Moore, 1978, Ann. S. Afr. Mus., 76 (4): 191-211.
Pseudobradya cornuta Lang, 1965 . Key to the species: Lang 1965.
P. crassipes Lang, 1965.
P. pectinifera Lang, 1965. This and the previous two species were described from off HMS in fine sand and detritus at about 26 m depth.
P. pulchera Lang, 1965. Described from rinsings of algae collected at DB.
P. lanceta Coull, 1986*. Sediments around eelgrass beds in WA and southern B.C..
Euterpina acutifrons (Dana, 1848). Planktonic, cosmopolitan between 66¡ N and 40¡ S. See Boxshall 1979, Bull. Brit. Mus. (Nat. Hist.). Zoology., 35(3): 201-264 redescription and distributional notes.
Key to the genera: Lang (1965), Huys et al. (1996).
Harpacticella sp. This asian brackish- and freshwater genus has been found in OR near the mouths of the Columbia and Klamath rivers.
Harpacticus compressus Frost, 1967* . Lower intertidal/shallow subtidal mud and sand with microalgae, epiphytes on eelgrass and macroalgae, epiphyton on docks. Described from Kodiak Island, Alaska.
H. compsonyx Monard, 1926*. A widely distributed warm-water species that was found by Kask (1982) on subtidal eelgrass in southern B.C..
H. pacificus Lang, 1965. Common in epiphytes on macroalgae, eegrass, and rocks. Part species group containing the european species H. obscurus, H. littoralis, and H. giesbrechti, which are difficult to separate. Given presence of other unidentified Pacific coast species of this group, care should be taken in assigning names.
H. septentrionalis Klie, 1939. Lower intertidal/shallow subtidal algae and eelgrass. Also in northern Europe and Japan.
H. spinulosus Lang, 1965. Described from rinsings of algae collected at DB. In Puget Sound and southern B.C., very abundant in sandy sediments around eelgrass and in unvegetated sand, where it is eaten by juvenile flatfish.
H. uniremis Kr¿yer, 1842. Mid intertidal to 20 m on a variety of habitats. Circumboreal distribution, occurring as far south as La Jolla, CA, where it occurred on subtidal algae (Gunnill 1982). From AK through WA, this species is abundant and forms a principal prey resource for outmigrating juvenile pink and chum salmon.
Perissocope biarticulatus Watkins, 1987. Described from CA in sandy sediments near macroalgae. Key to the species: Watkins 1987, J. Crust. Biol., 7 (2): 380-395.
Tigriopus californicus (Baker, 1912). Easily collected and recognized harpacticoid common all along the coast in supralittoral pools. Key to the species: Bradford 1967, Trans. r. Soc. N.Z. Zool., 10 (6): 51-59, It™ 1969, J. Fac. Sci. Hokkaido Univ., 17 (1): 58-77.
Z. aurelii Poppe, 1884 . Very common on eelgrass and macroalgae. It™ 1980, Publ. Seto mar. biol. Lab., 25 (1/4): 51-77 redescribed this species from Kodiak Island, AK. Lang (1948) and It™ (1980) considered this and Z. caeruleus Campbell, 1929 described from B.C., to be synonomous.
Z. robustus It™, 1974*. A japanese species reported by Kask et al. (1982) from dock epiphyton at the Pacific Biological Station, Nanaimo, B.C..
Z. spinatus hopkinsi Lang, 1965. Eelgrass, algae, and rock habitats from upper intertidal to shallow subtidal.
Zausodes septimus Lang 1965. Key to the species: Lang 1965.
Z. sextus Lang 1965. This and the previous species were described from off HMS at about 7 m depth.
Huntemannia jadensis Poppe, 1884. Common all along the coast in medium to coarse sand, from marine to tidal-fresh waters.
Nannopus palustris Brady, 1880. Estuaries, in brackish and freshwater mud and sand.
Arenolaophonte stygia Lang, 1965. Described from interstitial intertidal coarse sand near HMS. Watkins (1983) reported an apparently undescribed species collected from sediment trapped in Phyllospadix turf.
Echinolaophonte horrida (Norman, 1876). One specimen was found by Kask et al. (1982) on subtidal sand in southern B.C.. Key to the species: Lang 1965.
E. armiger briani Lang, 1965. A cosmopolitan species; holdfasts and epiphytes of algae and eelgrass.
Esola sp. A single specimen was collected near Little River, Mendocino County, CA by Watkins (1983).
Heterolaophonte discophora (Willey, 1929) . Macro- and microalgal and detritus-rich habitats. Boreal on both coasts of the U.S.; in the Pacific, occurs from AK to Monterey Bay and in Japan. Redescription: Lang 1965, It™ 1974, J. Fac. Sci. Hokkaido Univ., 19 (3): 546-640. Key to the species: Lang 1965. Species of the Ôquinquespinosa' group were removed to new genus Quinquelaophonte by Wells, et al. 1982, N.Z. J. Zool., 9: 151-184.
H. hamondi Hicks, 1975*. Common in intertidal eelgrass and sandy mud with detritus in Puget Sound; also southern B.C. (Kask et al. 1982).
H. longisetigera (Klie, 1950)*. Common in vegetated and detritus-rich habitats in Puget Sound and southern B.C.
H. mendax (Klie, 1939)*. One specimen found by Kask et al. (1982) in intertidal sandy mud at Nanaimo, B.C..
H. variabilis Lang 1965. Lower intertidal algae; very common on eelgrass epiphytes all along the coast.
Laophonte acutirostris Lang, 1965. Described from a tide pool containing coarse shell sand, near HMS.
Laophonte cornuta Phillipi, 1840 . Cosmopolitan; associated with algae and common all along the coast. Redescription: It™ 1968, J. Fac. Sci. Hokkaido Univ., 16 (3): 369-381.
L. elongata Boeck, 1872*. Specimens common in Puget Sound very similar to L. elongata triarticulata Monard, 1928. Watkins (1983) also found a single specimen of a species resembling but distinct from L. elongata in northern CA.
Laophonte sp. (inopinata group). Specimens similar to but distinct fromL. inopinata T. Scott, 1892 occur in tide pools near macroalgae (Watkins 1983) and also in epiphytes and microalgae in Puget Sound.
L. inornata A. Scott, 1902*. In southern B.C. and WA in intertidal and shallow subtidal sand in sheltered embayments.
Onychocamptus mohammed (Blanchard and Richard, 1891). Occurs over a wide geographical range and a variety of habitats. Intertidal and shallow subtidal in oligohaline and tidal-fresh regions of the Sacramento/San Joaquin delta and other estuaries along the Pacific coast, and in reservoirs of the Columbia River.
Paralaophonte asellopsiformis Lang, 1965. Described from a tide pool containing shell sand, near HMS. Lang (1965) remarked that this species was very similar to the mostly sand-dwelling genus Asellopsis, and described it as a "true sand-dweller". Key to the species: Lang 1965; Wells et al. 1982, N.Z. J. Zool., 9: 151-184.
P. congenera (Sars, 1908)*. Intertidal and shallow subtidal mud and sandy mud bays in WA and southern B.C..
P. pacifica Lang, 1965. Common from southern B.C. to southern CA in epiphytes and microalgae. A subspecies, P. pacifica galapagoensis Mielke 1981, Mikrofauna Meeresbodens, 52: 1-134. was described from the Galapagos Islands.
P. perplexa (T. Scott, 1898)*. A north Atlantic and Arctic species; commonly co-occurs with P. pacifica in southern B.C. and Puget Sound.
P. subterranea Lang, 1965. Described from interstices from sand near DB; also collected by Lang 1965 near HMS.
Paronychocamptus proprius Lang, 1965. Described from a tide pool with shell sand, stones, and algae near HMS.
P. sp. Another species of Paronychocamptus, similar to P. huntsmani (Willey, 1923) occurs near the mouth of the Columbia River.
Pseudonychocamptus paraproximus Lang 1965. Described from a Point Pinos, Monterey Bay tide pool, in fine sand. Common in Puget Sound shallow subtidal. Considered synonomous with P. proximus (Sars, 1908) by Mielke 1975, Mikrofauna Meeresbodens, 52: 1-134.. Key to the species: Lang 1965, Ceccherelli 1988, Vie Milieu, 38 (2): 155-171.
P. spinifer Lang 1965. Described from a tide pool with shell sand, stones, and algae near HMS. Common in Puget Sound around eelgrass and in lower intertidal and shallow subtidal sandy mud.
Quinquelaophonte capillata (C.B. Wilson, 1932)*. Mainly Atlantic species found by Kask et al. (1982) in intertidal sandy mud and shallow subtidal sand in southern B.C.; associated with eelgrass in Padilla Bay, northern WA. Redescription: Coull 1976, Trans. Am. Micros. Soc., 95 (1): 35-45.
Q. longifurcata Lang, 1965. Described from a tide pool with shell sand near HMS.
Key to the genera: Huys 1992.
Arenopontia dillonbeachia Lang 1965. Described from fine sand at about 3 m depth at DB; also found in Monterey Bay and in Japan by Ito 1969, J. Fac. Sci. Hokkaido Univ., 17 (1): 58-77 who described the male. Key to the species: Lang 1965; Bodiou & Colomines 1986, Crustaceana, 57 (3): 288-294.
Cerconeotes constrictus Lang 1965. Described as a species of Leptastacus from fine sand in a DB tide pool; also found year around in intertidal sandy mud in Nanaimo estuary, B.C. (Kask et al. 1982). Key to the species: Huys 1992, Med. Kon. Acad. Wetensch., Lett. Sch. Kunst. Belg., 54: 21-196.
Paraleptastacus spinicauda (T. and A. Scott, 1895). Reported from sand around eelgrass near the mouth of the Fraser river, B.C. by Webb 1983.
Sextonis incurvatus (Lang, 1965). Described as a species of Leptastacus from fine sand in Tomales Bay. Key to the species: Huys 1992, Med. Kon. Acad. Wetensch., Lett. Sch. Kunst. Belg., 54: 21-196.
Lourinia armatus (Claus, 1866). Geographically widespread in warm waters; found on algae near La Jolla, CA by Gunnill (1982). Watkins (1983) reported collecting a single specimen of another, undescribed species near Piedras Blancas Point, CA.
Normanella bolini Lang, 1965 . Key to the species: Lang 1965.
N. confluens Lang, 1965. This and the previous species were described from a tide pool with shell sand, stones, and algae near HMS.
Orthopsyllus linearis illgi (Chappuis, 1958). Described from Puget Sound, where it is abundant in intertidal and shallow subtidal sand beaches; redescribed by Lang (1965) from tide pools near HMS.
Key to the genera: Kunz 1981, Mitt. Zool. Mus. Univ. Kiel, 1 (8): 1-33.
Apodopsyllus vermiculiformis Lang, 1965 . Described from fine sand in a tide pool at Point Pinos, Monterey Bay. Key to the species: Lang 1965, Coull and Hogue 1978, Trans. am. micros. Soc., 97 (2): 149-159.
Scottopsyllus (Scottopsyllus) pararobertsoni Lang, 1965. Described from a tide pool with shell sand, stones, and algae near HMS. Key to the species: Lang 1965.
Parastenhelia hornelli Thompson and A. Scott, 1903*. Common in microalgae and epiphytes in a variety of intertidal and shallow subtidal habitats in southern B.C. and Puget Sound. Key to the species: Wells et al. 1982, N.Z. J. Zool., 9: 151-184.
P. spinosa (Fischer, 1860). A cosmopolitan species recorded in vegetated habitats from southern B.C. to La Jolla, CA.
Key to the genera: Huys et al. 1996. All members of this family are strongly dorsoventrally compressed and occur mainly on macroalgae and eelgrass blades.
Alteutha langi Monk, 1941 . Described from La Jolla, CA. Watkins (1983) found two other unidentified species from the CA intertidal.
Alteuthella sp. Watkins (1983) found an unidentified species near Piedras Blancas point.
Eupelte setacauda Monk, 1941 . Key to the species: Hicks 1982, South Africa. Zool. J. Linn. Soc., 75: 49-90.
E. simile (Monk, 1941). This and the previous species were described from La Jolla, CA. Watkins (1983) reported two (one as Paralteutha) apparently undescribed species from Mendocino County, CA intertidal.
Peltidium sp . Unidentified specimens have been found in coastal bays of WA, and reported from Mendocino County, CA intertidal by Watkins (1983).
This family is also dorsoventrally flattened.
Porcellidium fimbriatum Claus, 1863 . Found by Gunnill (1982) on algae near La Jolla. Kask et al. (1982) reported P. cf. fimbriatum from subtidal eelgrass and gravel in southern B.C..
P. viride (Phillipi, 1840). Puget Sound to La Jolla on eelgrass and macroalgae.
Rhizothrix sp.* Common in lower intertidal/shallow subtidal sandy habitats in Puget Sound. Key to the species: Bodin 1979, Vie Milieu, 27 (3-A): 311-357.
Key to the genera: Huys et al. 1996.
Geeopsis incisipes (Klie, 1913)*. A european species found in emergent marshes in Puget Sound estuaries.
Microarthridion littorale (Poppe, 1881) . Euryhaline; mud and muddy sand in estuaries and salt marshes; also occurs in epibenthic plankton. Probably a complex of several species.
Tachidius (Tachidius) discipes Giesbrecht, 1881. Northern european species common in oligohaline and tidal fresh water portions of estuaries in sand and mud, and in epibenthic plankton.
Tachidius (Neotachidius) triangularis Shen and Tai, 1963. One of the most abundant harpacticoids in marine-influenced tidal channels of coastal estuaries; also in eelgrass beds. Described from the Pearl River delta, South China, and may be introduced.
Family Tegastidae. All members of this family are strongly anterolaterally compressed, and have extremely hard integument. They are often associated with epibionts and colonial invertebrates Key to the genera: Huys et al. 1996.
Tegastes perforatus Lang 1965 . This and following species described from males only from about 7 m depth off HMS; Watkins 1983 figured female of this and females and males of three undescribed species in intertidal pools, and recorded a fourth undescribed species.
Syngastes serratus Lang, 1965.
Keys to the genera: Fiers 1995, Bull. Inst. R. Sci. Nat. Belg., Biol., 65: 151-236, Huys 1995, Hydrobiologia, 308 (1): 23-28.
Phyllopodopsyllus parabradyi Lang, 1965. Described from near HML in a tide pool with shell sand, stones, and algae. Key to the species: Lang 1965, Coull 1973, Proc. Biol. Soc. Wash., 86 (2): 9-24.
Phyllopodopsyllus borutzkyi Lang 1965. Described from tidepool algae, DB.
Paraschizopera trifida (Yeatman, 1980)*. Comb. nov. Huys, 1995, Hydrobiologia, 308 (1): 23-28, for Diagoniceps trifidus Yeatman, 1980. described from Puget Sound. Huys also gives a key to the species.
Key to the genera: Lang 1965.
Dactylopodella sp. Isolated specimens found in California (Watkins 1983) and Puget Sound. Key to the species: Hicks 1989, Nat. Mus. N.Z. Rec., 3 (10): 101-117.
Dactylopusia crassipes Lang, 1965 . Very abundant in estuaries where salinity is slightly lowered, and also in a variety of coastal vegetated and unvegetated habitats. Important prey for juvenile salmon.
D. glacialis (Sars, 1909). A North Atlantic and Arctic species; common but usually not abundant in microalgae and other vegetation.
D. paratisboides Lang , 1965. Microalgae and epiphytes on eelgrass and other substrates.
D. tisboides (Claus, 1863)*. A widely geographicall y distributed species (see Bodin 1964, Rec. Trav. Sta. Mar. Endoume, 51 (35) : 107-183) that has been found in southern B.C. and Puget Sound.
D. vulgaris inornata Lang, 1965. Abundant in epiphytes on eelgrass and macroalgae. Important in diets of juvenile salmon and flatfish.
Diarthrodes cystoecus Fahrenbach , 1954 . See Fahrenbach 1962 for a redescription and complete details of the biology of this algal leaf-mining species.
D. dissimilis Lang, 1965.
D. unisetosus Lang, 1965. This and the previous species were described from algae at DB, and were also found by Watkins (1983) in intertidal vegetated habitats.
Eudactylopus latipes typica Sewell, 1940. Eelgrass and macroalgae. Redescription: Lang 1965.
Idomene purpurocincta Norman and T. Scott, 1905 . European species found on algae and eelgrass. Supplemental description of specimens from California: Lang 1965.
Paradactylopodia serrata Lang, 1965 . mid- to lower intertidal on sand and muddy sand.
Parathalestris bulbiseta Lang, 1965 . Southern B.C. to La Jolla. Key to the species: Lang 1965.
P. californica Lang, 1965. This and previous species are usually associated with macroalgae, but can also occur on unvegetated sand.
P. jacksoni (T. Scott, 1898). A north Atlantic and Arctic species that also occurs in Alaska and southern B.C.
P. verrucosa Ito, 1970. A japanese species that was reported from epiphyton in southern B.C. by Kask et al. (1982).
Phyllothalestris mysis (Claus, 1863) . Algae rich lower intertidal habitats.
Rhynchothalestris helgolandica (Claus, 1863). Lower intertidal near algae; also occcurs in Puget Sound. Redescription: Huys 1990, Zool. J. Linn. Soc., 99: 51-115.
Thalestris longimana Claus, 1863. A european species that is abundant on algae in Puget Sound.
Scutellidium arthuri Poppe, 1884. On eelgrass and algae blades. Recorded from southern B.C. to southern CA, and in Japan. Redescription: Lang 1965. Key to the species: Branch 1975, Ann. S. Afr. Mus., 66 (10): 221-232.
S. hippolytes (Kroyer, 1863). Widely distributed on Atlantic coast in same habitats as above species. Recorded from southern B.C. to La Jolla.
Tisbe cf. furcata (Baird, 1837) . Eelgrass, detrital, and microalgal habitats. Due to many closely related 'sibling' species, identifications and distributions are difficult to establish for this genus. There are up to six unidentified/undescribed species in the NE pacific (Kask et al. 1982). T. cf. furcata appears to be cosmopolitan. It is an imortant prey item for juvenile salmon and other small fish.
Tachidiella parva Lang 1965. Described from off HMS in fine sand and detritus at about 26 m depth. This is a species incerta because here is a contradiction between the Lang's key and his diagnosis of the species : the key indicates 5 setae on the exopodite of the fifth leg, while the description and drawing mention only four.